The extreme, workerless inquilines.

[Teleutotje]

P.S. Merkur,

Those photographs from Saturday 16. January 2021 are new, better quality than those from the pdf....

[Teleutotje]

The extreme, workerless inquilines.

Teleutomyrmex Kutter, 1950

01) Teleutomyrmex schneideri Kutter, 1950
(= Tetramorium schneideri (Kutter, 1950))
(not Tetramorium schneideri Emery, 1898)
(= Tetramorium inquilinum Ward, Brady, Fisher & Schultz, 2015 ("2014"))
02) Teleutomyrmex kutteri Tinaut, 1990
(= Tetramorium kutteri (Tinaut, 1990))
(not Tetramorium semilaeve kutteri Santschi, 1927)
03) Teleutomyrmex seiferti Kiran & Karaman, in Kiran, et al. 2017
(= Tetramorium seiferti (Kiran & Karaman, in Kiran, et al. 2017))
04) Teleutomyrmex buschingeri Lapeva-Gjonova, in Kiran, et al. 2017
(= Tetramorium buschingeri (Lapeva-Gjonova, in Kiran, et al. 2017))

Not yet described species of extreme, workerless inquiline

05) The new, undescribed species from Teleutomyrmex from Farab, Turkmenistan… See Dlussky, et al. 1990.

Anergates Forel, 1874

06) Anergates atratulus (Schenck, 1852)
(= Myrmica atratula Schenck, 1852)
(= Tetramorium atratulum (Schenck, 1852))
[= Tomognathus atratulus (Schenck, 1852), obsolete combination.]
07) Anergates friedlandi Creighton, 1934
[= Tetramorium friedlandi ( Creighton, 1934)]

Tetramorium Mayr, 1855
(Only a few species in a big genus.)

08) Tetramorium microgyna Santschi, 1918
09) Tetramorium parasiticum Bolton, 1980

Pheidole Westwood, 1839
(Only a few species in a big genus.)

10) Pheidole neokohli Wilson, 1984
(= Pheidole kohli (Wasmann, 1915))
(= Anergatides kohli Wasmann, 1915)
(not Pheidole kohli Mayr, 1901)
11) Pheidole acutidens (Santschi, 1922)
(= Bruchomyrma acutidens Santschi, 1922)
12) Pheidole argentina (Bruch, 1932)
(= Gallardomyrma argentina Bruch, 1932)
13) Pheidole parasitica Wilson, 1984

Excluded from the extreme, workerless inquilines
Once this species was included in the extreme, workerless inquilines but now it is considered to be a workerless inquiline without extreme reductions, e.g. no pupoid males but normal ones. The decision to exclude it was made by Edward Osborne Wilson in 1984 in a study of the inquilines in the genus Pheidole.

Pheidole Westwood, 1839
(Only one species in a big genus.)

14) Pheidole kusnezovi Wilson, 2003
(= Pheidole symbiotica (Kusnezov, 1952))
(= Eriopheidole symbiotica Kusnezov, 1952)
(not Pheidole symbiotica Wasmann, 1909)

Distribution

01) Europe (Alps, Pyrenees and Northern Spain)
02) Europe (Southern Iberia)
03) Turkey (Anatolia)
04) Europe (Southern Balkans)

05) Turkmenistan

06) Europe
07) North America

08) Southern Africa
09) Southern Africa

10) Central Africa
11) South America
12) South America
13) India

14) South America

Host species

01), 02), 03), 04), 05), 06), 07), 08) and 09) Certain species of the genus Tetramorium Mayr, 1855

01) T. alpestre Steiner, Schlick-Steiner & Seifert, 2010 and T. impurum (Förster, 1850)
and maybe T. caespitum (Linnaeus, 1758)?
02) T. cf. caespitum (Linnaeus, 1758)
03) T. cf. chefketi Forel, 1911
04) T. cf. chefketi Forel, 1911

05) A species from the genus Tetramorium Mayr, 1855…

06) T. impurum (Förster, 1850), T. caespitum (Linnaeus, 1758), T. immigrans Santschi, 1927
and T. moravicum Kratochvil, 1941, T. diomedeum Emery, 1908, T. chefketi Forel, 1911
07) T. immigrans Santschi, 1927

08) T. sericeiventre Emery, 1877 and T. sepositum Santschi, 1918
09) T. avium Bolton, 1980

10), 11), 12), 13) and 14) Certain species of the genus Pheidole Westwood, 1839

10) P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912
11) P. nitidula Emery, 1888
12) P. nitidula Emery, 1888
13) P. indica Mayr, 1879

14) P. susannae Forel, 1886

A remark about synonymy

Anergates friedlandi Creighton, 1934 is now a synonym from Anergates atratulus (Schenck, 1852), more precisely an introduced form in North Americe. So, the name is Anergates atratulus (Schenck, 1852) or, if you follow Ward et al. 2015 ("2014"), Tetramorium atratulum (Schenck, 1852)...

Synonyms of host species

- T. caespitum (Linnaeus, 1758) (= Formica caespitum Linnaeus, 1758)
- T. impurum (Förster, 1850) (= Myrmica impura Förster, 1850)
- P. megacephala (Fabricius, 1793) (= Formica megecephala Fabricius, 1793)
- P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912 was originally described as P. punctulata Mayr, 1866 st. melancholica Santschi, 1912.
- P. susannae Forel, 1886 (= P. obscurior Forel, 1886)

[Teleutotje]

Yesterday, March 11, 2021, I was looking in the "Online Catalogue of the Ants of the World" on antwiki.org.
In it, the genus Teleutomyrmex stays like photograph 1, normal.
The tribe Teleutomyrmini stays like photograph 2. But what is that about Teleutomyrminae???

[Teleutotje]

Dumpert, K., 1978, “Das Sozialleben der Ameisen.” (First Edition.). Pareys Studientexte, nr. 18. Mit 95 Abbildungen. Verlag Paul Parey, Berlin und Hamburg, 1977. 253 pp.

Dumpert, K., 1994, “Das Sozialleben der Ameisen. 2., neubearbeitete Auflage.” (Second Edition.). Pareys Studientexte, nr. 18. Mit 94 Abbildungen. Verlag Paul Parey, Berlin und Hamburg, 1994. 257 pp.

99 references.

The reason why follows...

[Teleutotje]

Dumpert talks about Teleutomyrmex over a bit more than 2 pages and he includes a new, on Linsenmaier based, picture...
Picture 9.4 in edition 1, picture 7.4 in edition 2.

[Teleutotje]

Seifert 1996 about Teleutomyrmex...

[Teleutotje]

Seifert 2007 about Teleutomyrmex...

[Teleutotje]

Gösswald 1985 gave a discussion of almost 3 pages about Teleutomyrmex including a black and white version of the color picture of Linsenmaier and the next two pictures. In the determinationkey he included the text in picture 3.

[Teleutotje]

Kutter 1977, in his famous book of Swiss ants, says...

[Teleutotje]

Our dear Merkur, Professor Buschinger, in 2000, in Ameisenschutz aktuell, gives this...

[Teleutotje]

A few oldies about Anergates....

[Teleutotje]

An official literature list of Teleutomyrmex Kutter, 1950.

Baroni Urbani, C., 1967, “Le distribuzioni geografiche discontinue dei Formicidi mirmecobiotici.” Archivio Botanico e Biogeografico Italiano, vol. 43, [=(4)12],
p. 355-365.
Baur, A., Buschinger, A., Zimmermann, F. K., 1993, “Molecular cloning and sequencing of 18S rDNA gene fragments from six different ant species.” Insectes
Sociaux, vol. 40, p. 325-335.
Baur, A., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1995, “Mitochondrial DNA sequences reveal close relationships between social parasitic ants
and their host species.” Current Genetics, vol. 28, p. 242-247.
Baur, A., Sanetra, M., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1996, “Sequence comparisons of the internal transcribed spacer region of ribosomal
genes support close relationships between parasitic ants and their respective host species (Hymenoptera: Formicidae).” Insectes Sociaux, vol. 43, p. 53-67.
Bernard, F., 1967 ["1968"], “Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et
septentrionale.” Paris, Masson, 411 pp.
Blaimer, B. B., Ward, P. S., Schultz, T. R., Fisher, B. L., Brady, S. G., 2018, “Paleotropical Diversification Dominates the Evolution of the Hyperdiverse Ant Tribe
Crematogastrini (Hymenoptera: Formicidae).” Insect Systematics and Diversity, vol. 2, no. 5, art. 3, p. 1-14.
Bolton, B., 1976, “The ant tribe Tetramoriini (Hymenoptera: Formicidae). Constituent genera, review of smaller genera and revision of Triglyphothrix Forel.”
Bulletin of the British Museum (Natural History) (Entomology), vol. 34, p. 281-379.
Bolton, B., 1994, “Identification guide to the ant genera of the world.” Cambridge, Mass., Harvard University Press, 222 pp.
Bolton, B., 1995, “A taxonomic and zoogeographical census of the extant ant taxa (Hymenoptera: Formicidae).” Journal of Natural History, vol. 29, p. 1037-
1056.
Bolton, B., 1995, “A new general catalogue of the ants of the world.” Cambridge, Mass., Harvard University Press, 504 pp.
Bolton, B., 2003, “Synopsis and Classification of Formicidae.” Memoirs of the American Entomological Institute, vol. 71, 370 pp.
Borowiec, L., 2014, “Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae).” Genus (Wroclaw), vol. 25, no.
1-2, p. 1-340.
Bourke, A. F. G., Franks, N. R., 1991, “Alternative adaptations, sympatric speciation and the evolution of parasitic, inquiline ants.” Biological Journal of the
Linnean Society, vol. 43, no. 3, p. 157-178.
Brown, W. L., Jr., 1973, “A comparison of the Hylean and Congo-West African rain forest ant faunas.” P. 161-185 in: Meggers, B. J., Ayensu, E. S., Duckworth,
W. D. (ed.), 1973, “Tropical forest ecosystems in Africa and South America: a comparative review.” Smithsonian Institution Press, Washington, DC., viii + 350
pp.
Brun, R., 1952, “Das zentralnervensystem von Teleutomyrmex schneideri Kutt. (Hym. Formicid.). III. Mitteilung.” Mitteilungen der Schweizerischen
Entomologischen Gessellschaft, vol. 25, no. 2, p. 73-86.
Brun, R., 1959, “Le cerveau des fourmis et des insectes en général comme instrument de formation des réflexes conditionnés.” P. 11-25 in: International
Union of Biological Sciences, Union Internationale des Sciences Biologiques, Experimental Psychology and Animal Behaviour Section, 1959, “Animal
psychology seminars: Strasbourg University, October 1956, and Brussels University, August 1957.” London, Pergamon Press, 148 pp.
Buschinger, A., 1970, “Neue Vorstellungen zur Evolution des Sozialparasitismus und der Dulosis bei Ameisen (Hym., Formicidae).” Biologisches Zentralblatt,
vol. 88, no. 3, p. 273-299.
Buschinger, A., 1971, “Zur Verbreitung und Lebensweise sozialparasitischer Ameisen des Schweizer Wallis (Hym., Formicidae).” Zoologischer Anzeiger, vol.
186, no. 1/2, p. 47-59.
Buschinger, A., 1974, “Monogynie und Polygynie in Insektensozietäten.” P. 862-896 in: Schmidt, G. H. (ed.), 1974, “Sozialpolymorphismus bei Insekten.
Probleme der Kastenbildung im Tierreich.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, xxiv + 974 pp.
Buschinger, A., 1974, “Polymorphismus und Polyethismus sozialparasitischer Hymenopteren.” P. 897-934 in: Schmidt, G. H. (ed.), 1974,
“Sozialpolymorphismus bei Insekten. Probleme der Kastenbildung im Tierreich.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, xxiv + 974 pp.
Buschinger, A., 1985, “New records of rare parasitic ants (Hym., Form.) in the French Alps.” Insectes Sociaux, vol. 32, no. 3, p. 321-324.
Buschinger, A., 1986, “Evolution of social parasitism in ants.” Trends in Ecology and Evolution, vol. 1, no. 6, p. 155-160.
Buschinger, A., 1987, “Teleutomyrmex schneideri Kutter, 1950 and other parasitic ants found in the Pyrenees (Hymenoptera, Formicidae).” Spixiana, vol. 10,
no. 1, p. 81-83.
Buschinger, A., 1990, “Evolutionary transitions between types of social parasitism in ants, hypotheses and evidence.” P. 145-146 in: Veeresh, G. K., Mallik,
B., Viraktamath, C. A. (eds.), 1990, “Social insects and the environment. Proceedings of the 11th International Congress of IUSSI, 1990.” New Delhi, Oxford &
IBH Publishing Co., xxxi + 765 pp.
Buschinger, A., 1990, “Sympatric speciation and radiative evolution of socially parasitic ants. - Heretic hypotheses and their factual background.” Zeitschrift
für Zoologische Systematik und Evolutionsforschung, vol. 28, p. 241-260.
Buschinger, A., 1995, “Nicht am Ende: Die “Endameise” Teleutomyrmex schneideri.” Ameisenschutz aktuell, vol. 9, p. 1-7.
Buschinger, A., 1999, “Wiederfund der sozialparasitischen Ameise Teleutomyrmex schneideri in der Schweiz.” Mitteilungen der Schweizerischen
Entomologischen Gesellschaft, vol. 72, p. 277-279.
Buschinger, A., 2000, “Die “Endameise” Teleutomyrmex schneideri in der Schweiz: Erster Wiederfund nach 50 Jahren.” Ameisenschutz aktuell, vol. 14, p. 43-
45.
Buschinger, A., 2009, “Social parasitism among ants: a review (Hymenoptera: Formicidae).” Myrmecological News, vol. 12, p. 219-235.
Casevitz-Weulersse, J., Galkowski, C., 2009, “Liste actualisée des fourmis de France (Hymenoptera, Formicidae).” Bulletin de la Société Entomologique de
France, vol. 114, no. 4, p. 475-510.
Collingwood, C. A., 1956, “Ant hunting in France.” Entomologist, vol. 89, p. 105-108.
Collingwood, C. A., 1956, “A rare parasitic ant (Hym., Formicidae) in France.” Entomologist's Monthly Magazine, vol. 92, p. 197.
Csősz, S., Radchenko, A., Schulz, A., 2007, “Taxonomic revision of the Palaearctic Tetramorium chefketi species complex (Hymenoptera: Formicidae).”
Zootaxa, vol. 1405, p. 1-38.
Cuesta, D., García, F., García-Tejero, S. & Espadaler, X., 2009, “Resumen charla: Aportaciones a la biología de Teleutomyrmex schneideri Kutter, 1950: primer
caso de cría en cautividad. [Contributions to the biology of Teleutomyrmex schneideri Kutter, 1950: first case of keeping in captivity.]” [abstract].
Iberomyrmex, vol. 1, p. 24.
Cuesta-Segura, A.D., Espadaler, X., Garcia Garcia, F., 2017, “Hormigas de los brezales de Calluna cantábricos (NO España) (Hymenoptera: Formicidae). [Ants
of the Cantabrian Calluna-heathlands (NW Spain) (Hymenoptera: Formicidae).]” Iberomyrmex, vol. 9, p. 25-43.
Cuesta-Segura, A. D., García García, F., Catarineu, C., García-Tejero, S., Espadaler, X., 2018, “Actualización de la Distribución y Hospedadores de la Hormiga
Parasita Teleutomyrmex schneideri Kutter, 1950 en la Peninsula Ibérica (Hymenoptera: Formicidae).” Boletín de la Sociedad Entomológica Aragonesa
(S.E.A.), nº. 63 (31/12/2018), p. 235–239.
de la Mora, A., Sankovitz, M. & Purcell, J., 2020, "Ants (Hymenoptera: Formicidae) as host and intruder: recent advances and future directions in the study of
exploitative strategies." Myrmecological News, vol. 30, p. 53-71.
Dlussky, G. M., Fedoseeva, E. B., 1988 (“1987”), “[The origin and early stages of evolution of ants (Hymenoptera: Formicidae).]” P. 70-144 in: Ponomarenko,
A. G. (ed.), “[The Creaceous biocenotic crisis and the evolution of insects.]” [In Russian.]. Nauka, Moscow. 231 pp.
Dlussky, G. M., Soyunov, O. S., Zabelin, S. I., 1990 [“1989”], “Muravji Turkmenistana.”, or “[Ants of Turkmenistan.]” [In Russian.]. Ashkabad, Ylym Press, 275
pp.
Dobrzanska, J. & Dobrzanski, J., 1988, “Pasozytnictwo spoleczne u mrowek. Pasozytnictwo obowiazkowe.”, or “[Social parasitism in ants. Obligatory
parasitism.]” Kosmos, vol. 37, no. 4, p. 617-639.
-> Dobrzanska, J. & Dobrzanski, J., 1988, “1. Inkwilinizm.”, or “[Inquilinism.]” Kosmos, vol. 37, no. 4, p. 617-629.
-> Dobrzanska, J. & Dobrzanski, J., 1988, “2. Niewolnictwo.”, or “[Slavery.]” Kosmos, vol. 37, no. 4, p. 629-639.
Dumpert, K., 1978, “Das Sozialleben der Ameisen.” (First Edition.). Pareys Studientexte, nr. 18. Mit 95 Abbildungen. Verlag Paul Parey, Berlin und Hamburg,
1977, 253 pp.
Dumpert, K., 1994, “Das Sozialleben der Ameisen. 2., neubearbeitete Auflage.” (Second Edition.). Pareys Studientexte, nr. 18. Mit 94 Abbildungen. Verlag
Paul Parey, Berlin und Hamburg, 1994, 257 pp.
Espadaler, X., Cuesta, D., 2006, “Notas / Notes. Teleutomyrmex schneideri Kutter, 1950 en España (Hymenoptera, Formicidae).” Graellsia, vol. 62, no. 2, p.
261-262.
Espadaler, X., Gómez, K., 2014, “Tetramorium biskrense Forel, 1904 en España y Portugal peninsulares (Hymenoptera, Formicidae).” Boletín de la Sociedad
entomológica Aragonesa (SEA), vol. 55, p. 303-305.
García, F., 2019, “Parásitos y myrmecófilos. Hormigas parásitas sociales ibéricas.” LaMarabunta digital, Nueva Edición, Número 4, Junio 2019: p. 27-44.
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Kutter (Tribus Tetramorini).” Transactions of the IX-th International Congress of Entomology, Amsterdam, August 17-24 (1951), vol. 1, p. 446-448.
- His name is spelled “Göszwald” in this publication!
Gösswald, K., 1953, “Histologische Untersuchungen an der arbeiterlosen Ameise Teleutomyrmex schneideri Kutter (Hym. Formicidae).” Mitteilungen der
Schweizerischen Entomologischen Gessellschaft, vol. 26, no. 2, p. 81-128.
Gösswald, K., 1954, “Unsere Ameisen. I. Teil.” Stuttgart, Franckh'sche Verlagshandlung, 88 pp.
Gösswald, K., 1955, “Unsere Ameisen. II. Teil.” Stuttgart, Franckh'sche Verlagshandlung, 80 pp.
Gösswald, K., 1985, “Organisation und Leben der Ameisen.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, 355 pp.
Hölldobler, B. K., & Wilson, E. O., 1990, “The Ants.” Cambridge, Mass., Belknap Press of Harvard University Press, xiv + 732 pp.
Hölldobler, B. K., & Wilson, E. O., 1994, “Journey to the Ants: A Story of Scientific Exploration.” Cambridge, Mass., Belknap Press of Harvard University Press,
304 pp.
Kiran, K., Karaman, C., Lapeva-Gjonova, A. & Aksoy, V., 2017, “Two new species of the “ultimate“ parasitic ant genus Teleutomyrmex Kutter, 1950
(Hymenoptera: Formicidae) from the Western Palaearctic.“ Myrmecological News, vol. 25, p. 145-155.
Kusnezov, N., 1954, “Phyletische Bedeutung der Maxillar- und Labialtaster der Ameisen.” Zoologischer Anzeiger, vol. 153, no. 1/2, p. 28-38.
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2, p. 81-94.
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no. 3, p. 347-353.
Kutter, H., 1951, “Von dufttäuschenden Mörderinnen und berittener Königin.” Du: kulturelle Monatsschrift, vol. 11, no. 4, p. 44-47.
Kutter, H., 1952, “Über Plagiolepis xene Stärcke (Hym. Formicid.).” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 25, no. 2, p. 57-72.
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das Jahr 1969, vol. 171, (Ausgegeben am 31. Dezember 1968), p. 1-62.
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Formicidae).” European Journal of Entomology, vol. 97, no. 1, p. 95-117.
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Seifert, B., Buschinger, A., Aldawood, A., Antonova, V., Bharti, H., Borowiec, L., Dekoninck, W., Dubovikoff, D., Espadaler, X., Flegr, J. , Georgiadis, C., Heinze,
J., Neumeyer, R., Ødegaard, F., Oettler, J., Radchenko, A., Schultz, R., Sharaf, M., Trager, J., Vesnić, A., Wiezik, M., Zettel, H., 2016, “Banning paraphylies and
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Behavioural Ecology of Parasites.” CABI Publishing, Wallingford, UK and New York, USA, 369 pp.
Stumper, R., 1951, “Teleutomyrmex schneideri Kutter (Hym. Formicid.). II. Mitteilung. Über die Lebensweise der neuen Schmarotzerameise.” Mitteilungen
der Schweizerischen Entomologischen Gessellschaft, vol. 24, no. 2, p. 129-152.
Stumper, R., 1956, “Etudes myrmécologiques LXXVII. Les sécrétions attractives des reines de fourmis.” Mitteilungen der Schweizerischen Entomologischen
Gessellschaft, vol. 29, no. 4, p.373-380.
Stumper, R., Kutter, H., 1950, “Sur le stade ultime du parasitisme social chez les fourmis, atteint par Teleutomyrmex Schneideri (subtrib. nov.; gen. nov.;
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099 references.

[Teleutotje]

Now you can see what a wrong original can do with the result....

[Teleutotje]

Merkur wrote "Sorry, Teleutotje, I don't understand te context. Which original do you refer to?" My original was a pdf, my normal original is a word-document... Less problems with the text...

99 references in the included pdf...

[Teleutotje]

A new article, not much about Teleutomyrmex but it is mentioned as an endemic...

Biogeography of Iberian Ants (Hymenoptera: Formicidae).
by Alberto Tinaut and Francisca Ruano.
Diversity 2021, 13(2), 88, p. 1-25.
Published: 19 February 2021.

https://www.mdpi.com/1424-2818/13/2/88/htm
or
https://www.mdpi.com/1424-2818/13/2/88/pdf with two supplements.

or the blog "Biogeography of Iberian ants":

https://blog.myrmecologicalnews.org/202 ... rian-ants/

100 references!!!

[Teleutotje]

Something about this article. First the map in it where the Iberian Peninsula is divided in 8 regions:

Figure 1. Aprioristic refugium areas considered in the Iberian Peninsula. 1. Cantabric; 2. Pyrenean;
3. Mediterranean coast; 4. Atlantic; 5. Northern plateau; 6. Southern plateau; 7. Guadalquivir Valley;
8. Ebro Valley.

And then this table:

Table 5. List of 21 endemic species appearing in only one refugium area in the IP.

Pyrenean Area
Myrmicinae -
Myrmica lemasnei Bernard, 1967
Mediterranean Area
Leptanillinae -
Leptanilla theryi Forel, 1903
Formicinae -
Camponotus amaurus Espadaler, 1997
Cataglyphis gadeai De Haro & Collingwood, 2003
Cataglyphis humeya Tinaut, 1991
Myrmicinae -
Goniomma collingwoodi Espadaler, 1997
Goniomma compressisquama Tinaut, 1994
Messor timidus Espadaler, 1997
Teleutomyrmex kutteri Tinaut, 1990
Temnothorax ansei Catarineu, Barberá & Reyes-López, 2017
Temnothorax crepuscularis (Tinaut, 1994)
Northern Plateau Area
Leptanillinae -
Leptanilla charonea López, Martínez & Barandica, 1994
Myrmicinae -
Aphaenogaster ulibeli Gómez & Espadaler, 2018
Myrmoxenus bernardi (Espadaler, 1982)
Southern Plateau Area
Leptanillinae -
Leptanilla plutonia López, Martínez y Barandica, 1994
Myrmicinae -
Crematogaster fuentei Menozzi, 1922
Oxyopomyrmex magnus Salata & Borowiec, 2015
Guadalquivir Valley Area
Formicinae -
Cataglyphis floricola Tinaut, 1993
Cataglyphis tartessica Amor & Ortega, 2014
Myrmicinae -
Goniomma baeticum Reyes & Rodríguez, 1987
Ebro Valley Area
Myrmicinae -
Temnothorax caesari (Espadaler, 1997)

Of the 72 endemic species on 299 species on the Peninsula there are 21 species restricted to 1 area. And Teleutomyrmex kutteri is one of them!

[Teleutotje]

And a few quotes:

"On the other hand, although mountain ranges promoted species richness and endemicity, this pattern is not detectable for the Iberian and Palaearctic formicids. Only one ant species is exclusive to high mountains: Proformica longiseta (between 1800 and 3000 m a.s.l. in the Sierra Nevada mountains and the Baetic range). According to these findings, the majority of the Iberian ant endemic species are related with semiarid habitats with scarce vegetation at low altitudes, such as some of the species pertaining to the genus Temnothorax, as T. ansei, T. blascoi, T. caesari, T. crepuscularis. Another important number of endemic species are found in medium altitude mountain ranges, such as Teleutomyrmex kutteri (1700–2250 m a.s.l. in different Baetic ranges [106,107]), Temnothorax gredosi or T. conatensis (900–1500 m a.s.l.).
Whereas the rare species, i.e., those found in only one or two refugium areas, include for instance the social parasite ant species, where their way of life makes them difficult to find. These are known in very few and sometimes very distant areas, such as Rossomyrmex minuchae, Myrmoxenus bernardi or Anergates atratulus [112]. Another example of a rare species is Aphaenogaster cardenai, an Iberian endemic species of relatively wide distribution, but living in scarcely sampled habitats (the mesovoid shallow substratum, shallow caves and mines)."

and

"Alpine Species
Another important and more recent dispersal route is related with the Pleistocene glaciations, the last occurring 10,000 years ago. Many northern species survived and took refuge in the southern Palaearctic peninsulas, and during interglacial periods returned to the north and/or the populations climbed the slopes of the mountains. One good example of this groupis the pair of Iberian endemic species Formica frontalis/F. dusmeti, very close phylogenetically to F. truncorum, distributed in northern Europe (unpublished data). Probably the presence of T. kutterii in the south of the IP and T. schneideri in the northern IP and in the Alps is likely to have had this similar origin. Some species of Myrmica, Temnothorax, Lasius, etc, may belong to this group of alpine species, but this should be confirmed by phylogenetic molecular studies on the Mediterranean species."

and the references:

"106. Tinaut, A. Teleutomyrmex kutteri, spec. nov. A new species from Sierra Nevada (Granada, Spain). Spixiana 1990, 13, 201–208.
107. Reyes Lopez, J.; Benavente Martínez, A. Nueva cita de Teleutomyrmex kutteri Tinaut, 1990 (Hym. Formicidae) para la Península Ibérica. Bol. Soc. Entomol. Aragon. 2011, 49, 206.
112. Tinaut, A.; Ruano, F.; Martínez, M.D. Biology, distribution and taxonomic status of the parasitic ants of the Iberian Peninsula (Hymenoptera: Formicidae, Myrmicinae). Sociobiology 2005, 46, 1–42."

and a reference to look for:

"47. Sánchez-García, D.; Cuesta-Segura, D.A.; Trigos-Peral, G.; Arcos, J.; Catarineu, C.; García-García, F.; Herraiz, J.A.; Espadaler, X.; Gómez, K.; Tinaut, A. Checklist of the Iberian Myrmecofauna, In preparation."

[Teleutotje]

An article from Ameisenschutz Aktuell by Prof. Buschinger...

[Teleutotje]

First half of a new reference... Thought it already a long time but finally I found it again...

[Teleutotje]

And the rest...